A genetic history of the Balkans from Roman frontier to Slavic migrations

[crashdoc]

Anybody sent the links to the raw data for the new ancient samples and Serbs to Davidski yet? I figured somebody would've already done it but I might be suffering from bystander effect.

I just did (official genotypes from Reich site), about at the same time that you posted this message!

[crashdoc]

G25, courtesy of Davidski:

https://drive.google.com/file/d/1CmELrec...sp=sharing

[Michalis Moriakos]

Well done. We might ask him at some point if he can provide them for the East African adolescent. Even though he's very low coverage, I'd bet we'd at least get a good idea of what kind we're talking about: Nubian-like or Horner-like.

[Bale]

G25, courtesy of Davidski:

https://drive.google.com/file/d/1CmELrec...sp=sharing

be nice to see the North and East African outlier on G25

[rafc]

Thanks for the G25, I started digging through the samples:
Trogir Dragulin: 1-2nd centuries AD. Very diverse burial place. I26703 and I26705 match relatively well with IA samples from Croatia. Both seem to lack Anatolian ancestry. I26702 and I26708 are sibling, the boy being E-V13. They date from the later part of the period (say 150-200AD). They look very much like IA-Bulgarians, except that they have a bit of East Med admixture, which is mostly missing from IA-Bulgarians, but often present in the same profiles in the Roman era. I26704 and I26707 are both more East Med shifted. I26707 had rich grave goods, and looks fully Roman Imperial, being the most East Med shifted. So with only 6 samples we get three groups, something like IA Croatians, something like IA Bulgarians, and East-Med like people.

[rafc]

The Kormadin cemetery in Serbia is dated to 450-550AD, since it's classified as having Germanic grave goods it's probably related to the Ostrogoths, but again it's a very diverse bunch, with I27295 (female) showing more Germanic ancestry, and I27196 (R1a-Z94) more steppe-like roots. I27298 looks like an IA Croatian, while I27297 (E-V13) looks mostly like a Bulgarian IA with some additional Steppe ancestry, but other scenarios are possible.

[lukpl]

"At the fortified settlement of Brekinjova Kosa (Bojna, Croatia), two adult men dated to 770–890 cal CE were buried together in the same pit, the younger one (I26748) with healed skull trauma and with 97% Eastern European-related ancestry (...)"

Y-DNA: R-YP1337

My folk

[vasil]

I1116; 775-994 AD; Gomolava, Hrtkovci, Serbia; I2a1b-Y3120>S17250>Y5596>Z16971>Y5595>FT39645>Y12911>FT42350 (xFT43555,FT122179)

Y5596 level: Y5596/BY128+  A>C (9C)

Z16971 level: Y12910+  C>T (3T-1C)

Y5595 level: Y5595/Z16972+  G>T (1T)

FT39645 level: FT39645+  G>A (4A)

Y12911 level: Y12911/A2423+  C>T (3T)

FT42350 level: FT42350+  C>A (1A)

FT43555 level: FT43555-  G>C (1G)

FT122179 level: FT122179-  T>A (3T)

FT234871 level: *no calls*

Is this the earliest Dinaric sample found so far on the Balkans? I remember that some considered to be from the Bronze age are discounted as an archaeological mistake as they were not radiocarbon dated.

Yes.
The oldest I2a-Din was found in Turkey just south of Istanbul: Turkey_Byzantine_oEuropean__I10430 dated to 732 CE.
The second oldest are Pohansko samples (Czechia) from about 850 CE.
This Gomolava sample is dated to about 885 CE.

Thanks, I meant on the Balkans and it looks the earliest. However, this branch is now not common at all there, the area now in Serbia is dominated by Dinaric South, so probably there were further population movements.

PH908 is for sure already in place by this time. The Serb core in the early and high middle ages is in the south in Central Bosnia and Western Serbia/Rascia with a secondary expansion towards the SE(Kosovo/Shopluk), only after the Ottoman arrival with the rise of Moravian Serbia and later with the Great Serb migrations did it start to shift towards the north. This sample is likely not a Serb but related to the West Pannonian Slavs.

[lukpl]

Who is this guy? Goth?

Distance to: Croatia_Cepinski:I35081
0.02501696 Swedish_North
0.02862337 Swedish
0.03156009 Swedish
0.03156427 Norwegian
0.03219580 German_North-West
0.03273428 Norwegian_Bergen
0.03348995 Danish
0.03431551 Norwegian
0.03471427 German_Hamburg
0.03494706 Dutch_Central
0.03794174 Icelandic
0.03796646 Dutch
0.03905137 Frisian
0.03912989 German_West-Central
0.03980441 Slovak
0.04006956 UK_Ulster
0.04042237 German_East
0.04055380 German
0.04067736 Orcadian
0.04094662 Dutch_North
0.04205137 Scottish
0.04265833 Czech
0.04278582 Welsh
0.04299797 English
0.04321759 Shetlandic

and this
Distance to: Croatia_TrogirDobric:I26719
0.02921747 Norwegian_north
0.02982456 Danish
0.03008506 Shetlandic
0.03085733 Dutch_North
0.03138759 Norwegian
0.03205757 Orcadian
0.03223709 English
0.03224886 Welsh
0.03227250 Cornish
0.03254571 Swedish
0.03268026 Scottish
0.03280464 French_Finistere
0.03287465 Dutch
0.03337781 German_Hamburg
0.03359548 Icelandic
0.03382336 Irish
0.03401870 Breton_Cotes-d'Armor
0.03452384 Norwegian_Bergen
0.03490841 German_North-West
0.03578362 Breton_Finistere
0.03654786 Dutch_Central
0.03655371 Swedish
0.03698546 Belgian_Flemish
0.03744676 German
0.03819513 German_West-Central

[Southpaw]

This is for E-V13, some look Western-Balkan-like and more Eastern-Balkan-like, i do believe E-V13 might have been present where Urnfield spinoffs in Croatia, Northern Bosnia was present and cremation prevailed. Interesting, the Timacum E-V13 look more Western_Balkan-like, they were either Dardanian or Dalmatian based on grave classifications.

Target: Croatia_Nustar_E-V13:I28388
Distance: 2.2159% / 0.02215934 | ADC: 0.25x RC
54.8 Western_BalkansIA
19.0 AnatolianIA
17.4 East-BalkansIA
8.8 CentralBalkansIA

Target: Serbia_Kormadin:I27297_E-V13
Distance: 0.8817% / 0.00881680 | ADC: 0.25x RC
36.4 Western_BalkansIA
32.4 East-BalkansIA
23.2 CentralBalkansIA
8.0 AnatolianIA

Target: Croatia_TrogirDragulin_E-V13:I26702
Distance: 2.1629% / 0.02162870 | ADC: 0.25x RC
53.2 East-BalkansIA
16.8 Western_BalkansIA
15.2 AnatolianIA
14.8 CentralBalkansIA

Target: Serbia_TimacumKuline:I15537_E-V13
Distance: 2.3731% / 0.02373058 | ADC: 0.25x RC
51.8 Slavic
35.0 Western_BalkansIA
13.2 CentralBalkansIA

Target: Serbia_TimacumSlog:I15553_E-V13
Distance: 1.7015% / 0.01701503 | ADC: 0.25x RC
65.8 Western_BalkansIA
17.2 CentralBalkansIA
8.6 Slavic
8.4 AnatolianIA

Target: Serbia_TimacumSlog:I15544_E-V13
Distance: 1.7959% / 0.01795897 | ADC: 0.25x RC
53.0 Western_BalkansIA
24.6 CentralBalkansIA
22.4 East-BalkansIA

Target: Serbia_ViminaciumGrobalja_E-V13:I15526
Distance: 2.8967% / 0.02896694 | ADC: 0.25x RC
52.8 AnatolianIA
34.4 East-BalkansIA
12.8 CentralBalkansIA

Target: Serbia_ViminaciumGrobalja_E-V13:I15525
Distance: 2.4648% / 0.02464756 | ADC: 0.25x RC
74.4 AnatolianIA
25.6 Western_BalkansIA

Target: Serbia_ViminaciumGrobalja_E-V13:I15518
Distance: 2.9116% / 0.02911645 | ADC: 0.25x RC
63.2 East-BalkansIA
19.4 AnatolianIA
11.6 Western_BalkansIA
5.8 CentralBalkansIA

Target: Serbia_ViminaciumGrobalja_E-V13:I15513
Distance: 2.4279% / 0.02427886 | ADC: 0.25x RC
54.0 AnatolianIA
46.0 East-BalkansIA

Target: Serbia_ViminaciumRit:I15504_E-V13
Distance: 2.4408% / 0.02440847 | ADC: 0.25x RC
91.4 East-BalkansIA
5.2 Western_BalkansIA
3.4 CentralBalkansIA

Target: Serbia_ViminaciumRit:I15507_E-V13
Distance: 1.8659% / 0.01865906 | ADC: 0.25x RC
53.4 CentralBalkansIA
23.8 East-BalkansIA
9.4 Slavic
8.6 AnatolianIA
4.8 Western_BalkansIA

Target: Serbia_ViminaciumPirivoj:I15495_E-V13
Distance: 1.9723% / 0.01972318 | ADC: 0.25x RC
58.8 East-BalkansIA
26.2 Western_BalkansIA
15.0 AnatolianIA

Target: Serbia_ViminaciumPirivoj:I15490_E-V13
Distance: 2.7266% / 0.02726582 | ADC: 0.25x RC
68.2 East-BalkansIA
16.0 AnatolianIA
8.6 Western_BalkansIA
7.2 CentralBalkansIA

[rafc]

These are the Roman era samples (up to c. 600AD) on West-Eurasia PCA with the Y-haplogroups:

[Bukva_]

I1116; 775-994 AD; Gomolava, Hrtkovci, Serbia; I2a1b-Y3120>S17250>Y5596>Z16971>Y5595>FT39645>Y12911>FT42350 (xFT43555,FT122179)

Y5596 level: Y5596/BY128+  A>C (9C)

Z16971 level: Y12910+  C>T (3T-1C)

Y5595 level: Y5595/Z16972+  G>T (1T)

FT39645 level: FT39645+  G>A (4A)

Y12911 level: Y12911/A2423+  C>T (3T)

FT42350 level: FT42350+  C>A (1A)

FT43555 level: FT43555-  G>C (1G)

FT122179 level: FT122179-  T>A (3T)

FT234871 level: *no calls*

Is this the earliest Dinaric sample found so far on the Balkans? I remember that some considered to be from the Bronze age are discounted as an archaeological mistake as they were not radiocarbon dated.

Yes.
The oldest I2a-Din was found in Turkey just south of Istanbul: Turkey_Byzantine_oEuropean__I10430 dated to 732 CE.
The second oldest are Pohansko samples (Czechia) from about 850 CE.
This Gomolava sample is dated to about 885 CE.

Thanks, I meant on the Balkans and it looks the earliest. However, this branch is now not common at all there, the area now in Serbia is dominated by Dinaric South, so probably there were further population movements.

PH908 is for sure already in place by this time. The Serb core in the early and high middle ages is in the south in Central Bosnia and Western Serbia/Rascia with a secondary expansion towards the SE(Kosovo/Shopluk), only after the Ottoman arrival with the rise of Moravian Serbia and later with the Great Serb migrations did it start to shift towards the north. This sample is likely not a Serb but related to the West Pannonian Slavs.

I1116 is now my fathers closest match, much closer then any other ancient sample. Brekinjova Kosa samples doesn't fit his Slavic ancestry proxy as much as Bosnian sample I19561 or Krakauerberg samples. For context he has medieval ancestry from that area. Would like to hear ph2ter opinion on origin of kaikavian Slavs in medieval Slavonia.

[Capsian20]

This sample E-M123* maybe his origin is Near-Eastern ?
Target: Serbia_ViminaciumPirivoj:I15502
Distance: 2.5434% / 0.02543361
52.2 EEF
21.0 CHG
14.8 Iran_N
6.6 Steppe
5.4 Levant_Natufian_EpiP

Distance to: Serbia_ViminaciumPirivoj:I15502
0.03248373 Greek_Central_Anatolia:G2503
0.03326839 Greek_Cappadocia:658
0.03425961 Turkish_Trabzon:Trabzon21515
0.03485284 Greek_Cappadocia:662
0.03646141 Greek_Cappadocia:659
0.03667042 Turkish_Trabzon:TDP_Trabzon07
0.03702063 Greek_Cappadocia:663
0.03771996 Cypriot:CYP37
0.03776856 Turkish_Trabzon:TDP_Trabzon15
0.03807955 Greek_Trabzon:G25008
0.03823793 Greek_Dodecanese_RhodesOD2
0.03824747 Greek_Cappadocia:657
0.03864245 Greek_Cappadocia:661
0.03875663 Greek_Central_Anatolia:G2502
0.03902917 Greek_Central_Anatolia:G2501
0.03909005 Greek_Kos:GreeceKos4
0.03919444 Turkish_Trabzon:Trabzon21534
0.04040838 Cypriot:CYP30
0.04051742 Greek_Crete:760
0.04088749 Armenian_Urfa:Urfa004
0.04093298 Greek_Kos:GreeceKos10
0.04133312 Cypriot:CYP34
0.04136768 Greek_Kos:GreeceKos1
0.04141365 Turkish_Erzurum:TDP_Erzurum05
0.04173257 Greek_Trabzon:G25002

[Capsian20]

Even sample E-L791 is seems to me his origin is Near-Eastern

Distance to: Serbia_ViminaciumGrobalja:I15526
0.03098930 Cypriot:CYP32
0.03571987 Cypriot:CYP46
0.03664206 Cypriot:CYP45
0.03706779 Cypriot:CYP27
0.03741619 Cypriot:CYP19
0.03777792 Cypriot:CYP35
0.03801143 Greek_Dodecanese_RhodesOD4
0.03832027 Greek_Deep_Mani:ARE-19
0.03871410 Greek_Kos:GreeceKos7
0.03871998 Greek_Kos:GreeceKos5
0.03883570 Greek_Dodecanese_RhodesOD2
0.03897518 Cypriot:CYP30
0.03923664 Cypriot:CYP25
0.03943696 Greek_ApuliauG20
0.03963326 Greek_Kos:GreeceKos1
0.03981254 Greek_Crete:Crete7
0.04035379 Greek_Kos:GreeceKos8
0.04035609 Greek_Kos:GreeceKos4
0.04059313 Cypriot:CYP37
0.04092841 Greek_Dodecanese:668
0.04126344 Greek_Crete:764
0.04127046 Greek_Crete:776
0.04131853 Greek_Crete:795
0.04139630 Greek_Crete:B_Crete-2
0.04140802 Greek_Crete:730

Target: Serbia_ViminaciumGrobalja:I15526
Distance: 3.4667% / 0.03466723
62.8 EEF
14.4 Iran_N
9.6 CHG
8.2 Steppe
5.0 Levant_Natufian_contam

[Capsian20]

Well done. We might ask him at some point if he can provide them for the East African adolescent. Even though he's very low coverage, I'd bet we'd at least get a good idea of what kind we're talking about: Nubian-like or Horner-like.

And also sample North African please 
I32304