BTW MA-1 is dated 24000bp and Villabruna 1 is dated 14000bp so the expansion of ANE into the Pontic steppe happened smack dab in the middle of the LGM ie the Pontic steppe was inhabited for sure. Also Kostenki-12 is 30000 years old which then got replaced by EHG so I dont know if I would exactly call it a refugium but there was definitely continuous ocupation before, during and after the LGM.
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The time and place of origin of South Caucasian languages
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